International Code of Nomenclature

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International Code of Nomenclature for algae, fungi, and plants (Melbourne Code) adopted by the Eighteenth International Botanical Congress Melbourne, Australia, July 2011 prepared and edited by J. MCNEILL, Chairman F. R. BARRIE, W. R. BUCK, V. DEMOULIN, W. GREUTER, D. L. HAWKSWORTH, P. S. HERENDEEN, S. KNAPP, K. MARHOLD, J. PRADO, W. F. PRUD'HOMME VAN REINE, G. F. SMITH, J. H. WIERSEMA, Members N. J. TURLAND, Secretary of the Editorial Committee Koeltz Scientific Books 2012 The printed and only official version of the Code has been published as International Code of Nomenclature for algae, fungi, and plants (Melbourne Code). Regnum Vegetabile 154. Koeltz Scientific Books. ISBN 978-3-87429-425-6 © 2012, by International Association for Plant Taxonomy. Coding and design by Matúš Kempa, text to HTML by Dušan Senko and Matúš Kempa.

DIVISION I. PRINCIPLES Principle I: The nomenclature of algae, fungi, and plants is independent of zoological and bacteriological nomenclature. This Code applies equally to names of taxonomic groups treated as algae, fungi, or plants, whether or not these groups were originally so treated (see Pre. 8). Principle II: El uso de nombres de grupos taxonómicos está determinado por la existencia de tipos nomenclatoriales. The application of names of taxonomic groups is determined by means of nomenclatural types. Principle III: The nomenclature of a taxonomic group is based upon priority of publication. Principle IV: Each taxonomic group with a particular circumscription, position, and rank can bear only one correct name, the earliest that is in accordance with the rules, except in specified cases. Principle V: Scientific names of taxonomic groups are treated as Latin regardless of their derivation. Principle VI: The rules of nomenclature are retroactive unless expressly limited.

DIVISIÓN I. PRINCIPIOS La nomenclatura de algas, hongos, y plantas es independiente de la zoológica y bacteriológica. Este Código se aplica igualmente para nombres de grupos taxonómicos tratados como algas, hongos y plantas, independientemente de que originalmente se hayan tratado así o no. II. El uso de nombres de grupos taxonómicos está determinado por la existencia de tipos nomenclatoriales. III. La nomenclatura de un grupo taxonómico se basa en la prioridad de publicación. IV. Cada grupo taxonómico con una circunscripción, posición y rango particulares puede tener solamente un nombre correcto, el primero publicado y que cumple con las reglas, excepto en casos especificados. V. Los nombres científicos de los grupos taxonómicos se consideran en latín, sin importar su origen. VI. Las reglas de nomenclatura son retroactivas, excepto en casos notables.

Principios del Código Internacional de Nomenclatura Botánica (2012) V. Los nombres científicos de los grupos taxonómicos se consideran en latín, sin importar su origen. Ej. Ginkgo: Nombre derivado del japonés (pág. 811 de Kämpfer, Engelbert. 1712. Amoenitatum Exoticarum.)

Principios del Código Internacional de Nomenclatura Botánica (2012) V. Los nombres científicos de los grupos taxonómicos se consideran en latín, sin importar su origen. Género femenino: Ginkgo biloba Familia: Ginkgoaceae Orden: Ginkgoales Clase: Ginkgoopsida Chrysobalanus icaco L. Hidalgoa ternata La Llave Hildebrandtia africana Vatke Sequoia sempervirens (D. Don) Endl. Sequoiah (1770-1843), hijo de un mercader británico y una india cheroqui. Stephan L. Endlicher le dedicó el nombre genérico en 1847.

DIVISION II.  RULES AND RECOMMENDATIONS CHAPTER I.  TAXA AND THEIR RANKS Article 1 1.1. Taxonomic groups of any rank will, in this Code, be referred to as taxa (singular: taxon). 1.2. Fossil taxa (diatoms excepted) may be treated as morphotaxa. A morphotaxon is defined as a fossil taxon which, for nomenclatural purposes, comprises only the one part, life-history stage, or preservational state represented by the corresponding nomenclatural type. Note 1.  Any fossil taxon that is described as including more than one part, life-history stage, or preservational state is not a morphotaxon. Ex. 1. Alcicornopteris hallei J. Walton (in Ann. Bot, n.s., 13: 450. 1949) was described from fossil material that included a compression on the surface of a petrified nodule with anatomy permitting description of the rachides, sporangia, and spores of a pteridosperm. This species comprises two preservational stages, two life-history stages, and three parts of the plant and is therefore not a morphotaxon. Ex. 2.  Protofagacea allonensis Herend. & al. (in Int. J. Pl. Sci. 56: 94. 1995) was described on the basis of dichasia of staminate flowers, with anthers containing pollen grains, fruits, and cupules. This species comprises more than one part and more than one life-history stage and is therefore not a morphotaxon. 1.3. As in the case of form-taxa for asexual forms (anamorphs) of certain pleomorphic fungi (Art. 59), the provisions of this Code authorize the publication and use of names of morphotaxa (Art. 11.7). Article 2 2.1.  Every individual plant is treated as belonging to an indefinite number of taxa of consecutively subordinate rank, among which the rank of species (species) is basic. Article 3 3.1.  The principal ranks of taxa in descending sequence are: kingdom (regnum), division or phylum (divisio, phylum), class (classis), order (ordo), family (familia), genus (genus), and species (species). Thus, each species is assignable to a genus, each genus to a family, etc. Note 1.  Species and subdivisions of genera must be assigned to genera, and infraspecific taxa must be assigned to species, because their names are combinations (Art. 21.1, 23.1, and 24.1), but this provision does not preclude the placement of taxa as incertae sedis with regard to ranks higher than genus. Ex. 1.  The genus Haptanthus Goldberg & C. Nelson (in Syst. Bot. 14: 16. 1989) was originally described without being assigned to a family. Ex. 2.  The family assignment of the fossil genus Paradinandra Schönenberger & E. M. Friis (in Amer. J. Bot. 88: 467. 2001) was given as "incertae sedis". 3.2.  The principal ranks of nothotaxa (hybrid taxa) are nothogenus and nothospecies. These ranks are the same as genus and species. The prefix "notho" indicates the hybrid character (see App. I).

DIVISION II.  RULES AND RECOMMENDATIONS CHAPTER I.  TAXA AND THEIR RANKS Article 1 1.1. Taxonomic groups of any rank will, in this Code, be referred to as taxa (singular: taxon). Article 2 2.1.  Every individual plant is treated as belonging to an indefinite number of taxa of consecutively subordinate rank, among which the rank of species (species) is basic. Article 3 3.1.  The principal ranks of taxa in descending sequence are: kingdom (regnum), division or phylum (divisio, phylum), class (classis), order (ordo), family (familia), genus (genus), and species (species). Thus, each species is assignable to a genus, each genus to a family, etc.

Subdiv. Magnoliophytina (= Angiospermae) Article 2 2.1.  Every individual plant is treated as belonging to an indefinite number of taxa of consecutively subordinate rank, among which the rank of species (species) is basic. Ej. árbol de guanacaste Div. Spermatophyta Subdiv. Magnoliophytina (= Angiospermae) Cl. Magnoliopsida (= Dicotyledoneae) Subcl. Rosidae Ord. Fabales Fam. Fabaceae (= Leguminosae) Subfam. Mimosoideae Género Enterolobium especie cyclocarpum Enterolobium cyclocarpum (Jacq.) Griseb., Fl. Brit. W.I.: 226. 1860.

Article 6 6.1.  Effective publication is publication in accordance with Art. 29, 30, 31. 6.2.  Valid publication of names is publication in accordance with Art. 32, 33, 34, 35, 36, 37, 38, 39, 40, 41, 42, 43, 44, 45 or H.9 (see also Art. 61). Note 1.  For nomenclatural purposes, valid publication creates a name, and sometimes also an autonym (Art. 22.1 and 26.1), but does not itself imply any taxonomic circumscription beyond inclusion of the type of the name (Art. 7.1). 6.3.  In this Code, unless otherwise indicated, the word "name" means a name that has been validly published, whether it is legitimate or illegitimate (see Art. 12). Note 2.  When the same name, based on the same type, has been published independently at different times by different authors, then only the earliest of these "isonyms" has nomenclatural status. The name is always to be cited from its original place of valid publication, and later isonyms may be disregarded. Ex. 1.  Baker (Summary New Ferns: 9. 1892) and Christensen (Index Filic.: 44. 1905) independently published the name Alsophila kalbreyeri as a substitute for A. podophylla Baker (1891) non Hook. (1857). As published by Christensen, Alsophila kalbreyeri is a later isonym of A. kalbreyeri Baker, without nomenclatural status (see also Art. 33 Ex. 19). Ex. 2.  In publishing "Canarium pimela Leenh. nom. nov.", Leenhouts (in Blumea 9: 406. 1959) reused the illegitimate C. pimela K. D. Koenig (1805), attributing it to himself and basing it on the same type. He thereby created a later isonym without nomenclatural status. 6.4.  An illegitimate name is one that is designated as such in Art. 18.3, 19.5, or 52, 53, 54 (see also Art. 21 Note 1 and Art. 24 Note 2). A name which according to this Code was illegitimate when published cannot become legitimate later unless it is conserved or sanctioned. Ex. 3.  Anisothecium Mitt. (1869) when published included the previously designated type of Dicranella (Müll. Hal.) Schimp. (1856). When Dicranella was conserved with a different type, Anisothecium did not thereby become legitimate. Ex. 4.  Skeletonemopsis P. A. Sims (1995) was illegitimate when published because it included the original type of Skeletonema Grev. (1865). When Skeletonema was conserved with a different type, Skeletonemopsis nevertheless remained illegitimate and had to be conserved in order to be available for use. 6.5.  A legitimate name is one that is in accordance with the rules, i.e. one that is not illegitimate as defined in Art. 6.4. 6.6.  At the rank of family or below, the correct name of a taxon with a particular circumscription, position, and rank is the legitimate name which must be adopted for it under the rules (see Art. 11). Ex.5  The generic name Vexillifera Ducke (1922), based on the single species V. micranthera, is legitimate. The same is true of the generic name Dussia Krug & Urb. ex Taub. (1892), based on the single species D. martinicensis. Both generic names are correct when the genera are thought to be separate. Harms (in Repert. Spec. Nov. Regni Veg. 19: 291. 1924), however, united Vexillifera and Dussia in a single genus; the latter name is the correct one for the genus with this particular circumscription. The legitimate name Vexillifera may therefore be correct or incorrect according to different taxonomic concepts. 6.7.   The name of a taxon below the rank of genus, consisting of the name of a genus combined with one or two epithets, is termed a combination (see Art. 21, 23, and 24). Ex. 6.  Combinations: Mouriri subg. Pericrene, Arytera sect. Mischarytera, Gentiana lutea, Gentiana tenella var. occidentalis, Equisetum palustre var. americanum, Equisetum palustre f. fluitans. 6.8.  Autonyms are such names as can be established automatically under Art. 22.3 and 26.3, whether or not they appear in print in the publication in which they are created (see Art. 32.8, Rec. 22B.1 and 26B.1).

Article 6 6.1.  Effective publication is publication in accordance with Art. 29, 30, 31. 6.2.  Valid publication of names is publication in accordance with Art. 32, 33, 34, 35, 36, 37, 38, 39, 40, 41, 42, 43, 44, 45 or H.9 (see also Art. 61). 6.3.  In this Code, unless otherwise indicated, the word "name" means a name that has been validly published, whether it is legitimate or illegitimate (see Art. 12). Note 2.  When the same name, based on the same type, has been published independently at different times by different authors, then only the earliest of these "isonyms" has nomenclatural status. The name is always to be cited from its original place of valid publication, and later isonyms may be disregarded. 6.5.  A legitimate name is one that is in accordance with the rules, i.e. one that is not illegitimate as defined in Art. 6.4. 6.6.  At the rank of family or below, the correct name of a taxon with a particular circumscription, position, and rank is the legitimate name which must be adopted for it under the rules (see Art. 11). 6.7.   The name of a taxon below the rank of genus, consisting of the name of a genus combined with one or two epithets, is termed a combination (see Art. 21, 23, and 24).

Diagnosis latina Transcripción en latín de las características sobresalientes de una especie, que permiten distinguirla de otras especies afines. Tradicionalmente necesaria al describir especies nuevas. Ej.: Passiflora soliana sp. nov. La especie nueva es afín a Passiflora brevifila pero se distingue de ésta por diferentes caracteres florales: Flores más pequeñas (4–4.3 cm de diámetro), sin pedicelo o éste muy corto; brácteas florales más pequeñas (2–2.3 × 2–2.5 cm), ovadas a sub-orbiculares, con márgenes revolutos y connadas en su mitad proximal, con la superficie papilosa adaxialmente, sin un tomento marginal; corona de filamentos en dos series, los filamentos externos blancos y más cortos (7–9 mm). Species nova haec Passiflora brevifila Killip propinqua, sed taxon novum per sequentes floribus characteres aberratur: Flores breviores (4-4.3 cm diametro), sessiles (sine pedicello), bracteae florales breviores (2-2.3 x 2-2.5 cm diametro), ovatae usque ad suborbiculatas, cum marginibus revolutis et in parti proximali connatae, superficie adaxiali papillosa, sine tomento marginali, corona filamentosa biseriata, filamenta externa alba, breviora (7-9 mm longa).

Cyathea ursina (Maxon) Lellinger Diagnosis latina Cyathea x moralesiana A. Rojas, nothosp. nova, MES 2(3): 4-5. 2007 Difiere de Cyathea ursina (Maxon) Lellinger por frondas más largas, estípite más largo y muricado a espinoso, lámina más dividida y raquis menos densamente escamoso.   Nothospecies nova Cyathea ursina propinqua, sed frondibus longioribus, stipite longiore, muricato usque ad spinosum, lamina magis divisa et rachi laxiore squamoso dignoscenda. Cyathea multiflora Sm. Cyathea ursina (Maxon) Lellinger Cyathea x moralesiana A. Rojas

Diagnosis latina ¡Una diatomea! Amphipleura maya Céspedes-Vargas , nov. spec.   Valvae rhombicae usque ad rhombico-lanceolatas, 137-204 μm longae, 33-40 μm latae, 29-32 striae in 10 μm, cum apicibus leviter rotundatis. Costa axialis et raphe a A. lindheimeri similes; raphidis rami 38-47 µm longi. Differt a A. pellucida, A. lindheimeri et A. chiapasensis per latitudinem, strias transapicales et raphidis ramos. Valvas rómbico a rómbico-lanceoladas con extremos poco redondeados. Costilla axial y rafe como en A. lindheimeri. Longitud 137-204 µm, ancho 33-40 µm, estrías 29-32/ 10 µm. Longitud de la fisura del rafe 38-47 µm, ocupando entre un 1/3 a 1/4 del eje apical. El ancho es la principal característica que diferencia A. maya de las especies, A. pellucida, A. lindheimeri y A. chiapasensis, las cuales son muy similares a la especie (Tabla 1). Estrías transapicales discrepan con las especies antes mencionadas. Al igual que la longitud de la fisura del rafe.

Descripción de una especie nueva: Datos obligatorios 1) Nombre de la especie. 2) Indicación clara de que se trata de una especie nueva: sp. nov., spec. nova, especie nueva, new species… 3) Diagnosis latina de caracteres diagnósticos A partir de 2012 puede ser una diagnosis en lengua inglesa 4) Datos del espécimen tipo: Lugar de recolecta, nombre del recolector, número de recolecta, fecha 5) Ubicación física del espécimen tipo en un herbario, un museo o una colección privada

Una nueva especie de Fevillea (Cucurbitaceae: Zanonieae) de Costa Rica A. Estrada & D. Santamaría. J. Bot. Res. Inst. Texas 4(1): 45-49. 2010 Fevillea narae A. Estrada & D. Santamaría, sp. nov. (Figs. 1, 2). Tipo: COSTA RICA: San José: Cantón de Tarrazú, Distrito San Lorenzo, cuenca del Naranjo y Paquita, Quebrada Salitrillo, a 1 km del cruce a San Isidro y Cerro Nara, 9°28'40"N, 84°02'00"W, 300–400 m, 2 mar 2005 (fls. pistiladas), D. Santamaría & J.F. Morales 821 (holotipo: CR; isotipos: INB, MO). Species nova a omnibus speciebus generis Fevilleae per sequentes characteres distinguenda: Folia integra, glabra, laminae apex glandularis, sed petiolus, basis et margenes eglandulares, flores masculi solitarii vel inflorescentiis sub-umbelliformibus, brevibus et paucifloribus dispositi.

Una nueva especie de Fevillea (Cucurbitaceae: Zanonieae ) de Costa Rica Armando Estrada Ch. Daniel Santamaría A. Museo Nacional de Costa Rica Instituto Nacional de Biodiversidad Apartado Postal 749-1000 Apdo. 22-3100 San José, COSTA RICA Santo Domingo de Heredia, COSTA RICA aestrada@museocostarica.go.cr dsantamaria@inbio.ac.cr Resumen: Fevillea narae, una nueva especie procedente del cantón de Tarrazú, al sur de San José, en la vertiente pacífica de la cordillera de Talamanca, Costa Rica, es descrita e ilustrada. Esta nueva especie representa la segunda especie de Fevillea para la región mesoamericana, junto a Fevillea cordifolia. Abstract: Fevillea narae, a new species from the cantón of Tarrazú along the pacific slope of cordillera Talamanca, Costa Rica, south of San José is described and illustrated. This species is the second species of Fevillea for the Mesoamerican region. Fevillea (Cucurbitaceae) es un género neotropical, compuesto por lianas o bejucos dioicos y representado por 7 especies (Robinson y Wunderlin 2005; Monro 2009). Estas especies crecen usualmente a orillas de ríos, bordes de bosques, claros de vegetación y orillas de caminos y carreteras. El género se caracteriza por presentar hojas con glándulas; cáliz con escamas glandulares (excepto F. passiflora); pétalos con un apéndice o cresta adaxial sobre la costilla media que se extiende desde la base hasta la mitad del pétalo; flores estaminadas con 5 estambres, separados y anteras biloculares; frutos grandes, usualmente con una cicatriz circunferencial hacia el ápice (excepto F. passiflora) y semillas grandes (Robinson y Wunderlin 2005; Monro 2009). La gran mayoría de especies del género son suramericanas; sólo Fevillea cordifolia, una especie ampliamente distribuida en todo el neotrópico, es conocida en la región de Mesoamérica y El Caribe (Jeffrey 2001; Robinson y Wunderlin 2005). Fevillea narae constituye la segunda especie de este género para la región y por el momento la primera especie del grupo con una distribución restringida a esta zona. La nueva especie es hasta ahora endémica de Costa Rica y se localiza al noroeste de la Cord. de Talamanca, en la vertiente pacífica central del país. Ésta es una zona de elevaciones medias, con bosques muy húmedos y nublados, una topografía abrupta con gran cantidad de ríos y quebradas y un pronunciado gradiente altitudinal, condiciones que favorecen una alta diversidad biológica, que actualmente presenta problemas serios de conservación por deforestación y alteración de los hábitats naturales. La región hasta hace poco ha empezado a explorarse más exhaustiva y sistemáticamente, dando como resultado el descubrimiento de varias especies nuevas como Chamaedorea piscifolia Hodel, G. Herrera & Cascante, Lacmellea zamorae J.F. Morales, Epidendrum montis-narae Pupulin & L. Sánchez S., Macroclinium montis-narae Pupulin, Guarea constricta Al. Rodr.; Ruyschia moralesii Hammel; Ruellia norvegigratiosa McDade & E. Tripp. Fevillea narae A. Estrada & D. Santamaría, sp. nov. (Figs. 1, 2). Tipo: COSTA RICA: San José: Cantón de Tarrazú, Distrito San Lorenzo, cuenca del Naranjo y Paquita, Quebrada Salitrillo, a 1km del cruce a San Isidro y Cerro Nara, 9°28'40"N, 84°02'00"W, 300–400 m, 2 Mar 2005 (fls. pistiladas), D. Santamaría & J.F. Morales 821 (holotipo: CR; isotipos: INB, MO). Species nova a omnibus speciebus generis Fevilleae per sequentes characteres distinguenda: Folia integra, glabra, laminae apex glandularis, sed petiolus, basis et margenes eglandulares, flores masculi solitarii vel inflorescentiis sub-umbelliformibus, brevibus et paucifloribus dispositi. Esta nueva especie difiere del resto de especies del género Fevillea por sus hojas enteras, glabras, con glándulas en el ápice de la hoja, pero ausentes en pecíolos, base o márgenes de la lámina; además, por sus flores estaminadas solitarias o en inflorescencias subumbeliformes cortas y poco floreadas. J. Bot. Res. Inst. Texas 4(1): 45 – 49. 2010

Descripción de una especie nueva El protólogo: Conjunto de datos de la publicación original de una especie de plantas, necesarios para tipificar e identificar la especie (diagnosis, descripción, hábitat, distribución geográfica, discusión, ejemplares citados, ilustración) Drymonia tomentulifera Kriebel, sp. nova Tipo: Costa Rica. Alajuela: Cuenca del Sarapiquí, Virgen del Socorro, 10º15'25"N 84º10'20"W, 800 m, 21 de julio 2002, R. Kriebel 536 (Holotipo: INB; isotipos: CR, MO). Fig. 1. A Drymonia pilifera Wiehler affinis, sed indumento tomentuloso, corola minore, maculata, cum superficie externa sericea differt.

Descripción de una especie nueva Byttneria osaënsis Cristóbal, sp. nova Holotipo: Costa Rica. Puntarenas, cantón de Osa, Península de Osa, Reserva Forestal Golfo Dulce, entrada a Chocuaco, Bahía Chal, 8º43'20”N, 83º26'30”W, 15 nov 1992 (fr), Reinaldo Aguilar 1468 (CR). Isotipos: CTES, MO. Figs. 1 y 2 Planta scandens inermis, foliis integribus, subcoriaceis, rigidis, elipticis, lamina ad basim rotundata vel subcordata; nervi medii basis cum nectario nigrescenti, prominenti, instructo, 5-6 mm longo; hypophyllo heterotricho, pilis stellatis, diminutis, in areolis hialinis, sed supra venis rufis; petala glabra, cum lamina lanceolata; fructus complanatus, lignosus, coccis ca. 4 cm longis et 3,6 cm latis, castaneo-rubescentibus, aculeis piramidalibus, 2-4 mm longis, sparsis, acutis, striatis, pubescentibus, pilis diminutis, simplicibus.

Descripción de una especie nueva Trichopilia x ramonensis J. García & Mora-Ret. ex C.O. Morales, nothosp. nova Holotipo: Alajuela; cantón San Ramón, distrito San Rafael, Berlín, 10°03’N, 84°28’O. Leg. Luis Acosta s.n.; floreció en cultivo en marzo de 1991. USJ-57879 (en líquido). Fig. 1. Nothospecies nova hic descripta, sed multos annos ab egregiis orchideologis nota, apud J. García et Mora-Retana in anno 1992 iam citata sed adhuc non descripta. Epiphyta parva, pseudobulbis dense aggregatis, elongatis, 6-11 cm longis, 1,7-2,1 cm latis, plus minusve rectangulatis, complanatis, interdum falcatis. Foliis elliptico-lanceolatis, petiolo 0,5-2,0 cm longo, lamina 12-24 cm longa, 3,5-6,9 cm lata. Inflorescentia brevis, saepe 1 vel 3 (5) floribus. Sepalis petalisque rubriusculis, sepalis lateralibus ad basim 1,0-1,5 cm connatis; labello roseo-purpureo, inter Trichopiliam marginatam et T. suavem intermedie structurato, 6,1-7,5 cm longo, 4,7-5,8 cm lato [complanato disposito], conspicue venato, ad apicem ampliato, profunde emarginato et obscure tetralobato, marginibus valde undulatis simul irregulatis; columna erecta, 2,8-3,1 cm longa, ad basim 1,0-1,5 cm cum labello connata; clinandrio supra anthera conspicue laciniato. Fructum non vidi.

Descripción de una especie nueva 4 cm 3 mm 2 mm Fig. 1. Trichopilia x ramonensis. A. Hábito; planta con restos de dos inflorescencias. B. Sépalo dorsal. C. Sépalos laterales. D. Pétalo. E. Labelo (algo extendido y aplanado). F. Columna con pedicelo (N.B. clinandrio incompleto). G. Antera, vista dorsal. H. Antera, vista ventral. I. Polinios. A: Dibujo del autor; planta del Jardín Botánico Lankester. B-I: Dibujos del holotipo (USJ), por Marcia González Garay. 2 cm

Descripción de una especie nueva Cedrela ngobe Köcke, T.D.Penn. & Muellner, sp. nov. A speciebus aliis foliis 6–7-jugis, infra breviter puberulis, petalis 6–6.5 mm longis, capsula 4–6 cm longa differt. – Type: Panama, Comarca Ngöbe-Buglé, foothills of Cerro Guánico, A.V. Köcke , O. Cáceres , H. Wessels & M. Piepenbring 180409/02 (holo K; iso FR, PMA, UCH). Fig. 1 . EDINBURGH J O U R N A L O F B O T A N Y 72 ( 2 ): 225 – 233 (2015) © Trustees of the Royal Botanic Garden Edinburgh (2015) doi:10.1017/S0960428615000098 CEDRELA NGOBE (MELIACEAE), A NEW SPECIES FROM PANAMA AND COSTA RICA A.V. KÖCKE, A.N. MUELLNER-RIEHL , O. CÁCERES & T.D. PENNINGTON FIG. 1. Cedrela ngobe Köcke, T.D.Penn. & Muellner. A, leaf and inflorescence; B, detail of leaf undersurface; C, flower; D, cross section of a male flower; E, capsules; F, seed (A–D, Köcke et al . 180409/02; E–F, Köcke et al. 180409/01).

Protólogo de Licania hedbergii (Chrysobalanaceae)

Protólogo de Burmeistera intii Gómez Laur. & L. D Protólogo de Burmeistera intii Gómez Laur. & L.D. Gómez (Campanulaceae)